It is clear from the photo how a beetle collects and deposits pollen grains among Othonna coronopifolia disc florets, using six feet while fully focussed on eating. A bit like a hiker collecting and losing dust particles from his boots on a mountain trail, totally oblivious of the nano-events he is causing.
Ray florets may be variously curled back in the same flowerhead, usually strongly at night. Ray and disc florets are both bisexual.
The stems in picture differ in colour from white to brown with some added maculation (Manning 2007; Manning and Goldblatt, 1997; Bond and Goldblatt, 1984; iNaturalist; JSTOR).